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Watters, G. T. (1992). Unionids, fishes, and the species-area curve. Journal of Biogeography. 19:481-490.

The number of fish species in thirty-seven systems of the Ohio River may be predicted by the area of the drainage basin. On a large river system scale, the number of unionid species is directly related to the number of fish species present and, to a lesser extent, the drainage area. In small systems, such as headwater tributaries, the number of unionid species is related to the drainage basin area. Unionid diversity in systems intermediate in size may be related to both. This may be the result of transient fishes in small streams depositing glochidia. The overall distribution and diversity of unionids within the study area is dependent upon the distribution and diversity of their fish hosts.

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